JPH02503988A - 改変雄蕊細胞を有する植物 - Google Patents
改変雄蕊細胞を有する植物Info
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Abstract
Description
Claims (20)
- 1.外来DNA配列、好ましくは外来キメラDNA配列により形質転換される核 ゲノムを有する植物の細胞であって、前記外来DNA配列が (a)前記植物の雄蕊細胞に産生されまたは過剰産生された際に雄蕊細胞の代謝 、機能および/または発育を顕著に阻害する第1RNA、蛋白もしくはポリペプ チドをコードする雄不稔性DNA、および (b)前記雄不稔性DNAの発現を植物の雄蕊細胞、好ましくは葯、花粉および /または花糸細胞、特にタペートおよび/または葯の表皮細胞にて選択的に指示 しうる第1プロモータからなり、前記雄不稔性DNAは前記第1プロモータと同 じ転写単位に位置しかつその制御下にあることを特徴とする植物の細胞。
- 2.前記外来DNA配列が、好ましくは前記雄不稔性DNAと同じ遺伝子部位に 、 (c)植物の少なくとも特定組織または少なくとも特定細胞に存在する際に植物 を少なくとも前記特定組織もしくは特定細胞に第2RNA、蛋白もしくはポリペ プチドを含有しない他の植物から容易に分離しうるようにする前記第2RNA、 蛋白もしくはポリペプチドをコードする標識DNA、および(d)前記標識DN Aの発現を少なくとも前記特定組織もしくは特定細胞にて指令しうる第2プロモ ータをさらに含み、前記標識DNAは前記第2プロモータと同じ転写単位に位置 しかつその制御下にある請求の範囲第1項記載の細胞。
- 3.前記外来DNA配列が、 (e)前記第1蛋白もしくはポリペプチドを前記雄蕊細胞の葉緑体もしくはミト コンドリア中に導入しうるトランジットペプチドをコードし、前記雄不稔性DN Aおよび前記第1プロモータと同じ転写単位に位置すると共に前記雄不稔性DN Aと前記第1プロモータとの間に位置する第1DNA、および/または(f)第 2蛋白もしくはポリペプチドを少なくとも前記特定組織もしくは特定細胞の葉緑 体もしくはミトコンドリア中に導入しうるトランジットペプチドをコードし、前 記標織DNAおよび前記第2プロモータと同じ転写単位に存在すると共に前記標 識DNAと前記第2プロモータとの間に位置する第2DNAをさらに含む請求の 範囲第1項または第2項記載の細胞。
- 4.前記雄不稔性DNAがRNアーゼ、特にRNアーゼTIも∬ しくはバルナーゼ;DNアーゼ、特にエンドヌクレアーゼ、殊にEcoRI;プ ロテアーゼ、殊にパパイン」、特にパパインチモーゲンもしくはパパイン活性蛋 白;グルカナーゼ;リパーゼ、特にホスホリパーA2;脂質パルオキシダーゼ; 細胞壁インヒビター;細胞トキシン;またはリポチーム、特にT^A29遺伝子 、TA26遺伝子もしくはTA13遺伝子によりコードされるmRNAに対する リボチームをコードするか、またはアンチセンスDNA、特にTA29遺伝子、 TA26遺伝子もしくはTA13遺伝子のアンチセンスDNAである請求の範囲 第1項〜第3項のいずれか一項に記載の細胞。
- 5.前記雄不稔性DNAが、植物ホルモンの合成を触媒する酵素、特にAgro bacteriumT−DNAの遺伝子1、遺伝子2もしくは遺伝子4によりコ ードされる酵素をコードする請求の範囲第1項〜第4項のいずれか一項に記載の 細胞。
- 6.前記標識DNAが除草剤耐性遺伝子、特にsfrもしくははsfrV遺伝子 ;除草剤に対しより低い親和性を有する除草剤用の改質標的酵素、特にグリホセ ートの標的としての改変5−エノールピルビルスキメート−3−ホスフェートシ ンターゼまたはグルタミンシンセターゼ阻止剤(たとえばホスフィノトリシン) の標的としての改変グルタミンシンセターゼをコードする遺伝子;少なくとも前 記特定組織もしくは特定細胞に色を与える蛋白もしくはポリペプチをコードする 遺伝子、特に遺伝子A1もしくはGUS遺伝子;前記植物にストレス耐性を付与 する蛋白もしくはポリペプチドをコードする遺伝子、特にMn−スーパーオキシ ドジスムターゼをコードする遺伝子;病気もしくは害虫耐性を付与する蛋白もし くはポリペプチドをコードする遺伝子、特にBacillusthurngie nsisのンドトキシンをコードして昆虫耐性を付与する遺伝子;または殺細菌 住ペプチドをコードして細菌耐性を付与する遺伝子である請求の範囲第2項〜第 5項のいずれか一項に記載の細胞。
- 7.前記第1プロモータがPTA29,PTA26,PTA13またはTA29 ,TA26もしくはTA13にハイブリッド化しうるタペート特異性mRNAを コードするDNAのプロモータである請求の範囲第1項〜第6項のいずれか一項 に記載の細胞。
- 8.前記第2プロモータが構成プロモータ、特に35Sプロモータ、35S′3 プロモータ、PNOSプロモータもしくはPOCSプロモータ;傷誘発性プロモ ータ、特にTR1もしくはTR2プロモータ;光合成活性を有する植物組織にて 遺伝子発現を選択的に指令するプロモータ、特にSSUプロモータ;または葉細 胞、花弁細胞もしくは種子細胞、特に種子被覆細胞にて遺伝子発現を選択的に指 命ずるプロモータである請求の範囲第2項〜第7項のいずれか一項に記載の細胞 。
- 9.植物、特にAgrobacteriumで感染しうる植物の細胞を形質転換 させるのに適し、請求の範囲第1項〜第8項のいずれか一項に記載の外来PNA 配列、特にpTTM4′pTTM6′pTTM6A−,pTTM8,pTVEP 1,pTVEP2,pTVE62もしくはpTVE63を含むベクター。
- 10.細胞の核ゲノムに安定的に組込まれた請求の範囲第2項〜第8項のいずれ か一項に記載の外来DNA配列を有し、これにより前記雄不稔性DNAが植物の 雄蕊細胞にて選択的に発現されて前記雄蕊細胞に前記第1DNA、蛋白もしくは ポリペプチドを生成させると共に、前記標識DNAを前記植物の少なくとも特定 組織もしくは特定細胞にて発現させて前記植物を未形質転換植物から分離しうる ようにする雄不稔性植物、およびこの横物の繁殖材料の生産方法において、(a )前記外来DNA配列を前記細胞の核ゲノム中に導入することにより前記植物の 細胞を形質転換させ、次いで(b)前記植物および繁殖材料を前記細胞から再生 させる非生物学的工程を特徴とする堆不稔性植物および二の植物の繁殖材料の生 産方法。
- 11.請求の範囲第三項〜第8項のいずれか一項に記載のー植物細胞を含有する 植物細胞培養物。
- 12.請求の範囲第1項〜第8項のいずれか一項に記載の植物細胞を含有する植 物、特にトウモロコシ、馬鈴薯、トマト、菜種、アルファルファ、ヒマワリ、綿 、セロリ、玉ネギ、クローバー、大豆、タバコ、油菜野菜もしくは甜菜。
- 13.請求の範囲第12項記載の植物の種子。
- 14.(a)(i)前記第2プロモータと前記標識DNAとの両者を含み、特に 前記標識DNAが除草剤に対する耐性を付与し、特に前記雄不稔性植物の細胞に おける核ゲノムに安定的に組込まれたsfrもしくはsfrV遺伝子である請求 の範囲第2項〜第8項のいずれか一項に記載の外来DNA配列を含有する雄不稔 性植物を、(ii)前記標識DNAを持たない、特に前記除草剤耐性を付与する 前記標識DNAを持たないホモ接合性の雄稔性植物と他家受粉させ、次いで(b )前記雄稔性植物の少なくとも前記特定組織もしくは特定細胞にて前記標識DN Aが発現しないことを利用して、前記雄稔性植物を前記他の雄不稔性植物から分 離する工程を特徴とするハイブリッド種子の生産方法。
- 15.前記雄不稔性植物が細胞の核ゲノム中に安定的に組込まれた少なくとも2 種の異なる標識DNAを含有し、かつ前記雄稔性植物が前記2種の標識DNAの 一方を含有するが他方を含有せず、さらに前記雄不稔性植物の少なくとも前記特 定組織もしくは特定細胞にて前記他方の標識DNAが発現しないことを利用して 前記雄稔性植物を前記雄不稔性植物から分離し、前記他方の標識DNAが好まし くは除草剤に対する耐性を付与する請求の範囲第14項記載のハイブリツド種子 の生産方法。
- 16.請求の範囲第14項または第15項記載の方法により得られるハイブリッ ド種子。
- 17.請求の範囲第16項記載のハイブリッド種子を成長させて得られるハイブ リッド植物。
- 18.請求の範囲第7項記載の第1プロモータ、特にpTA29、pTA26も しくはpTA13。
- 19.前記雄不稔性DNAが第1プロモータの制御下で天然には存在せず、かつ /または前記標識DNAと同じ遺伝子部位には天然に存在しない請求の範囲第1 項〜第8項のいずれか一項に記載の外来キメラDNA配列。
- 20.核ゲノムに安定的に組込まれかつそこでRNA、蛋白もしくはポリペプチ ドとして発現しうる外来遺伝子物質を含んだ植物およびこの植物のたとえば種子 のような繁殖材料を生産するに際し、(a)前記RNA、蛋白もしくはポリペプ チドを発現しない出発植物細胞もしくは植物組織から前記外来遺伝子物質を含む 形質転換された植物細胞もしくは植物組織を生産し、(b)再生された植物また はこれら植物の繁殖材料またはその両者を前記外来遺伝子物質を含む前記形質転 換された植物細胞もしくは植物組織から生産し、かつ(c)必要に応じ前記再生 された植物もしくは繁殖材料またはその両者を生物学的に複製する非生物学的工 程からな生産方法において、前記外来遺伝子物質を含む前記形質転換された植物 細胞もしくは植物組織を生産する工程は、前記出発植物細胞もしくは植物組織の 核ゲノムを請求の範囲第1項〜第8項のいずれか一項に記載の外来DNA配列、 並びに前記植物細胞もしくは植物組織における前記外来DNA配列の発現を可能 にする制御要素により形質転換させて、形質転換された植物細胞もしくは植物組 織、並びに後代全体にわたりそれらから生産された植物および繁殖材料にて前記 外来DNA配列の安定な組込みを生ぜしめることを特徴とする植物およびその繁 殖材料の生産方法。
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1988
- 1988-04-28 GB GB888810120A patent/GB8810120D0/en active Pending
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1989
- 1989-04-26 AT AT96107004T patent/ATE395425T1/de not_active IP Right Cessation
- 1989-04-26 EP EP89401194A patent/EP0344029B1/en not_active Expired - Lifetime
- 1989-04-26 ES ES89401194T patent/ES2097745T3/es not_active Expired - Lifetime
- 1989-04-26 AT AT89401194T patent/ATE148498T1/de not_active IP Right Cessation
- 1989-04-26 DE DE68927721T patent/DE68927721T2/de not_active Expired - Lifetime
- 1989-04-26 DE DE68929558T patent/DE68929558D1/de not_active Expired - Lifetime
- 1989-04-26 EP EP96107004A patent/EP0737749B1/en not_active Expired - Lifetime
- 1989-04-27 IE IE138689A patent/IE81154B1/en not_active IP Right Cessation
- 1989-04-27 CA CA000597953A patent/CA1340324C/en not_active Expired - Lifetime
- 1989-04-27 WO PCT/EP1989/000495 patent/WO1989010396A1/en unknown
- 1989-04-27 IL IL9009589A patent/IL90095A/en active IP Right Revival
- 1989-04-27 HU HU763/89A patent/HU217413B/hu unknown
- 1989-04-27 AU AU35371/89A patent/AU621113C/en not_active Expired
- 1989-04-27 CA CA000616878A patent/CA1341494C/en not_active Expired - Lifetime
- 1989-04-27 ZA ZA893136A patent/ZA893136B/xx unknown
- 1989-04-27 JP JP1504514A patent/JP3020530B2/ja not_active Expired - Lifetime
- 1989-04-28 PT PT90420A patent/PT90420B/pt not_active IP Right Cessation
- 1989-12-27 DK DK198906684A patent/DK175585B1/da not_active IP Right Cessation
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1993
- 1993-03-05 US US08/027,580 patent/US6372967B1/en not_active Expired - Lifetime
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1994
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1995
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1996
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1997
- 1997-04-22 GR GR970400885T patent/GR3023224T3/el unknown
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1998
- 1998-06-27 HK HK98107204A patent/HK1008051A1/xx not_active IP Right Cessation
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1999
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Cited By (2)
Publication number | Priority date | Publication date | Assignee | Title |
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JPH04504355A (ja) * | 1989-02-02 | 1992-08-06 | パラディン ハイブリッズ インコーポレーテッド | ハイブリッド種子生産の分子的方法 |
JP2007060979A (ja) * | 2005-08-31 | 2007-03-15 | Asahi Breweries Ltd | システインプロテアーゼ遺伝子を導入した葯の裂開を抑制する植物 |
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