ID29922A - Produk multi-enzim dengan aktivitas glukoamilase, proteolitik dan ksilanase dan metode untuk memproduksinya dengan cara fermentasi kondisi padat dari kulit gandum dengan aspergillus niger - Google Patents

Produk multi-enzim dengan aktivitas glukoamilase, proteolitik dan ksilanase dan metode untuk memproduksinya dengan cara fermentasi kondisi padat dari kulit gandum dengan aspergillus niger

Info

Publication number
ID29922A
ID29922A IDW00200101818A ID20011818A ID29922A ID 29922 A ID29922 A ID 29922A ID W00200101818 A IDW00200101818 A ID W00200101818A ID 20011818 A ID20011818 A ID 20011818A ID 29922 A ID29922 A ID 29922A
Authority
ID
Indonesia
Prior art keywords
aspergillus niger
activities
glukoamilase
cocilanase
proteolitic
Prior art date
Application number
IDW00200101818A
Other languages
English (en)
Indonesian (id)
Inventor
Pierre Jean Labeille
Jean-Luc Alan Guy Baret
Francis Lucien Duchiron
Original Assignee
Gie Agro Ind
Priority date (The priority date is an assumption and is not a legal conclusion. Google has not performed a legal analysis and makes no representation as to the accuracy of the date listed.)
Filing date
Publication date
Family has litigation
First worldwide family litigation filed litigation Critical https://patents.darts-ip.com/?family=9541183&utm_source=***_patent&utm_medium=platform_link&utm_campaign=public_patent_search&patent=ID29922(A) "Global patent litigation dataset” by Darts-ip is licensed under a Creative Commons Attribution 4.0 International License.
Application filed by Gie Agro Ind filed Critical Gie Agro Ind
Publication of ID29922A publication Critical patent/ID29922A/id

Links

Classifications

    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2405—Glucanases
    • C12N9/2408—Glucanases acting on alpha -1,4-glucosidic bonds
    • C12N9/2411—Amylases
    • C12N9/2428—Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N1/00—Microorganisms, e.g. protozoa; Compositions thereof; Processes of propagating, maintaining or preserving microorganisms or compositions thereof; Processes of preparing or isolating a composition containing a microorganism; Culture media therefor
    • C12N1/14—Fungi; Culture media therefor
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/24—Hydrolases (3) acting on glycosyl compounds (3.2)
    • C12N9/2402—Hydrolases (3) acting on glycosyl compounds (3.2) hydrolysing O- and S- glycosyl compounds (3.2.1)
    • C12N9/2477—Hemicellulases not provided in a preceding group
    • C12N9/248—Xylanases
    • C12N9/2482—Endo-1,4-beta-xylanase (3.2.1.8)
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12N—MICROORGANISMS OR ENZYMES; COMPOSITIONS THEREOF; PROPAGATING, PRESERVING, OR MAINTAINING MICROORGANISMS; MUTATION OR GENETIC ENGINEERING; CULTURE MEDIA
    • C12N9/00—Enzymes; Proenzymes; Compositions thereof; Processes for preparing, activating, inhibiting, separating or purifying enzymes
    • C12N9/14—Hydrolases (3)
    • C12N9/48—Hydrolases (3) acting on peptide bonds (3.4)
    • C12N9/50—Proteinases, e.g. Endopeptidases (3.4.21-3.4.25)
    • C12N9/58—Proteinases, e.g. Endopeptidases (3.4.21-3.4.25) derived from fungi
    • C12N9/62—Proteinases, e.g. Endopeptidases (3.4.21-3.4.25) derived from fungi from Aspergillus
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01003—Glucan 1,4-alpha-glucosidase (3.2.1.3), i.e. glucoamylase
    • C—CHEMISTRY; METALLURGY
    • C12—BIOCHEMISTRY; BEER; SPIRITS; WINE; VINEGAR; MICROBIOLOGY; ENZYMOLOGY; MUTATION OR GENETIC ENGINEERING
    • C12Y—ENZYMES
    • C12Y302/00—Hydrolases acting on glycosyl compounds, i.e. glycosylases (3.2)
    • C12Y302/01—Glycosidases, i.e. enzymes hydrolysing O- and S-glycosyl compounds (3.2.1)
    • C12Y302/01008—Endo-1,4-beta-xylanase (3.2.1.8)
    • Y—GENERAL TAGGING OF NEW TECHNOLOGICAL DEVELOPMENTS; GENERAL TAGGING OF CROSS-SECTIONAL TECHNOLOGIES SPANNING OVER SEVERAL SECTIONS OF THE IPC; TECHNICAL SUBJECTS COVERED BY FORMER USPC CROSS-REFERENCE ART COLLECTIONS [XRACs] AND DIGESTS
    • Y02—TECHNOLOGIES OR APPLICATIONS FOR MITIGATION OR ADAPTATION AGAINST CLIMATE CHANGE
    • Y02E—REDUCTION OF GREENHOUSE GAS [GHG] EMISSIONS, RELATED TO ENERGY GENERATION, TRANSMISSION OR DISTRIBUTION
    • Y02E50/00—Technologies for the production of fuel of non-fossil origin
    • Y02E50/10—Biofuels, e.g. bio-diesel

Landscapes

  • Life Sciences & Earth Sciences (AREA)
  • Chemical & Material Sciences (AREA)
  • Health & Medical Sciences (AREA)
  • Organic Chemistry (AREA)
  • Engineering & Computer Science (AREA)
  • Zoology (AREA)
  • Bioinformatics & Cheminformatics (AREA)
  • Genetics & Genomics (AREA)
  • Wood Science & Technology (AREA)
  • Biochemistry (AREA)
  • General Health & Medical Sciences (AREA)
  • General Engineering & Computer Science (AREA)
  • Biotechnology (AREA)
  • Medicinal Chemistry (AREA)
  • Microbiology (AREA)
  • Biomedical Technology (AREA)
  • Molecular Biology (AREA)
  • Mycology (AREA)
  • Botany (AREA)
  • Tropical Medicine & Parasitology (AREA)
  • Virology (AREA)
  • Preparation Of Compounds By Using Micro-Organisms (AREA)
  • Enzymes And Modification Thereof (AREA)
  • Micro-Organisms Or Cultivation Processes Thereof (AREA)
  • Fodder In General (AREA)
  • Cereal-Derived Products (AREA)
IDW00200101818A 1999-01-25 2000-01-25 Produk multi-enzim dengan aktivitas glukoamilase, proteolitik dan ksilanase dan metode untuk memproduksinya dengan cara fermentasi kondisi padat dari kulit gandum dengan aspergillus niger ID29922A (id)

Applications Claiming Priority (1)

Application Number Priority Date Filing Date Title
FR9900775A FR2788782B1 (fr) 1999-01-25 1999-01-25 Produit multienzymatique a activites glucoamylasique, proteolytique et xylanasique et procede pour sa production par fermentation a l'etat solide de son de ble avec aspergillus niger

Publications (1)

Publication Number Publication Date
ID29922A true ID29922A (id) 2001-10-25

Family

ID=9541183

Family Applications (1)

Application Number Title Priority Date Filing Date
IDW00200101818A ID29922A (id) 1999-01-25 2000-01-25 Produk multi-enzim dengan aktivitas glukoamilase, proteolitik dan ksilanase dan metode untuk memproduksinya dengan cara fermentasi kondisi padat dari kulit gandum dengan aspergillus niger

Country Status (22)

Country Link
US (1) US6667066B2 (pt)
EP (1) EP1022329B1 (pt)
JP (1) JP2002534973A (pt)
CN (1) CN1203170C (pt)
AT (1) ATE294852T1 (pt)
AU (1) AU775240B2 (pt)
BR (1) BRPI0007600B1 (pt)
CA (1) CA2361264C (pt)
DE (1) DE60019825T2 (pt)
DK (1) DK1022329T3 (pt)
ES (1) ES2244395T3 (pt)
FR (1) FR2788782B1 (pt)
HU (1) HUP0104907A2 (pt)
ID (1) ID29922A (pt)
MA (1) MA25276A1 (pt)
MX (1) MXPA01007528A (pt)
PL (1) PL201768B1 (pt)
PT (1) PT1022329E (pt)
RU (1) RU2246536C2 (pt)
TR (1) TR200102156T2 (pt)
WO (1) WO2000043496A1 (pt)
ZA (1) ZA200107009B (pt)

Families Citing this family (48)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
WO2002088342A1 (en) * 2001-04-30 2002-11-07 Biocon India Limited An enzyme preparation for improved baking quality and a process for preparing the same
BR0312121A (pt) * 2002-06-14 2007-04-03 Diversa Corp xinalases, ácidos nucléicos codificando-as e métodos para preparar e empregá-las
US20040191362A1 (en) * 2003-03-26 2004-09-30 Dasappa Indrani Synergistic improver mix
EP1622471A4 (en) * 2003-05-12 2007-02-14 Alltech Inc METHODS FOR IMPROVING THE NUTRITIONAL QUALITY OF RESIDUES IN THE ENERGY PRODUCTS INDUSTRY, ALCOHOLIC BEVERAGES, HUMAN AND ANIMAL FEEDING
MXPA06000067A (es) 2003-07-02 2007-01-25 Diversa Corp Glucanasas, acidos nucleicos que las codifican y metodos para hacerlas y usarlas.
EP3219806B1 (en) * 2004-03-25 2020-05-06 Novozymes, Inc. Methods for degrading or converting plant cell wall polysaccharides
BRPI0509718B1 (pt) * 2004-04-09 2015-12-01 Asahi Breweries Ltd Processo de fabricação de koji líquido e processo de fabricação de um alimento ou drinque fermentado”
KR101222413B1 (ko) * 2004-05-27 2013-01-16 다니스코 유에스 인크. 아스퍼질러스 가와치 산-안정성 알파 아밀라제의 이종 발현및 입상 전분 가수분해에서의 이용
RU2409659C2 (ru) * 2005-07-22 2011-01-20 Асахи Брюэриз, Лтд. Способ получения жидкого коджи
MY143774A (en) * 2005-09-20 2011-07-15 Asahi Breweries Ltd Method of producing liquid koji having enhanced plant fiber degradation enzyme, liquid koji obtained by the method and use thereof
AU2006298348B8 (en) * 2005-10-05 2012-02-02 Asahi Breweries, Ltd. Method of producing filamentous fungus culture product
JP4096009B2 (ja) * 2005-10-12 2008-06-04 アサヒビール株式会社 組換えタンパクの製造方法
CN100448979C (zh) * 2005-12-09 2009-01-07 山东省中协食品添加剂研究开发中心 一种酶法生产木糖的方法
FR2898605B1 (fr) * 2006-03-17 2008-06-27 J Soufflet Sa Ets Complement nutritionnel pour milieu de saccharification-fermentation dans la production d'ethanol
US8304212B2 (en) * 2006-07-10 2012-11-06 Dyadic International, Inc. Methods and compositions for degradation of lignocellulosic material
CN101117628B (zh) * 2006-08-01 2010-12-08 中国农业科学院北京畜牧兽医研究所 生产植酸酶的方法
AU2007356171B8 (en) 2006-08-04 2014-01-16 Bp Corporation North America Inc. Glucanases, nucleic acids encoding them, and methods for making and using them
BRPI0717181B1 (pt) 2006-09-21 2022-04-05 Basf Enzymes Llc Ácido nucleico, cassete de expressão, vetor, veículo de clonagem, microrganismo transgênico, polipeptídeo de fitase, polipeptídeo imobilizado ou ácido nucleico imobilizado, processos de produção de um polipeptídeo recombinante, para hidrolisar um inositol-hexafosfato em inositol e fosfato inorgânico, para degomagem de óleo, para produção de um alimento, produto alimentício ou suplemento alimentar, para processamento de grãos de milho e de sorgo, alimento, produto alimentício ou suplemento alimentar, composição, e, uso de um polipeptídeo
MX2009003034A (es) 2006-09-21 2009-11-18 Verenium Corp Fosfolipasas, acidos nucleicos que las codifican, y metodos de hacerlas y usarlas.
PL2069490T5 (pl) 2006-12-21 2018-09-28 Basf Enzymes Llc Amylazy i glukoamylazy, kwasy nukleinowe kodujące te związki oraz sposoby wytwarzania tych związków oraz stosowania ich
CA2723113C (en) * 2008-04-29 2018-06-26 Icm, Inc. Pretreatment of grain slurry with alpha-amylase and a hemicellulase blend prior to liquefaction
EP2362737B1 (en) * 2008-10-24 2012-12-05 Unilever PLC, a company registered in England and Wales under company no. 41424 Frozen confection
CN102292437B (zh) 2008-12-19 2015-06-24 杜邦营养生物科学有限公司 生产酶产品的方法
NZ596459A (en) 2009-05-21 2013-11-29 Verenium Corp Phytases, nucleic acids encoding them and methods for making and using them
UA111708C2 (uk) 2009-10-16 2016-06-10 Бандж Ойлз, Інк. Спосіб рафінування олії
UA109884C2 (uk) 2009-10-16 2015-10-26 Поліпептид, що має активність ферменту фосфатидилінозитол-специфічної фосфоліпази с, нуклеїнова кислота, що його кодує, та спосіб його виробництва і застосування
FR2960551B1 (fr) * 2010-05-27 2012-09-14 Comptoir Agricole Production de molecules d'interet en milieu solide
CN102550895A (zh) * 2010-12-28 2012-07-11 雀巢公司 来自于食物固态发酵的酶制剂
CN103492579A (zh) * 2011-04-29 2014-01-01 丹尼斯科美国公司 用纤维素酶和葡糖淀粉酶提高发酵制备乙醇的产率
CN102676483A (zh) * 2012-01-15 2012-09-19 河南科技大学 一种一菌多酶菌株产蛋白酶的方法
MD4186C1 (ro) * 2012-02-20 2013-06-30 Институт Микробиологии И Биотехнологии Академии Наук Молдовы Tulpină de fungi Fusarium gibbosum - producătoare de proteaze acide şi neutre, xilanaze şi b-glucozidaze
WO2013163703A1 (pt) * 2012-05-03 2013-11-07 Petróleo Brasileiro S.A. - Petrobras Processo integrado de produção de formulações enzimáticas a partir de resíduos agroindustriais e produção de biocombustíveis
MD4285C1 (ro) * 2013-02-28 2014-12-31 Институт Микробиологии И Биотехнологии Академии Наук Молдовы Tulpină de fungi Trichoderma koningii Oudemans - producătoare de proteaze acide, neutre şi alcaline
CN103305428B (zh) * 2013-06-26 2015-02-18 武汉合缘绿色生物工程有限公司 一种黑曲霉菌株及其应用
FR3016768B1 (fr) * 2014-01-27 2016-02-26 J Soufflet Ets Utilisation d'une composition enzymatique dans l'alimentation des ruminants
WO2017155803A1 (en) 2016-03-08 2017-09-14 Basf Enzymes Llc Methods for using phytase in ethanol production
CN106754838B (zh) * 2017-01-20 2020-06-09 上海绿农饲料科技有限公司 一种提高黑曲霉产蛋白酶能力的方法
EP3530743A1 (en) 2018-02-21 2019-08-28 Cambridge Glycoscience Ltd Method of production
BR112021002910A2 (pt) 2018-08-15 2021-07-20 Cambridge Glycoscience Ltd novas composições, seu uso e métodos para sua formação
CN110373401A (zh) * 2019-07-22 2019-10-25 武汉新华扬生物股份有限公司 一种麻花用复配酶制剂及其制备方法和应用
JP2022545650A (ja) 2019-08-16 2022-10-28 ケンブリッジ グリコサイエンス エルティーディー バイオマスを処理してオリゴ糖および関連組成物を生成する方法
CN115279208A (zh) 2019-12-12 2022-11-01 剑桥糖质科学有限公司 低糖多相食品
CN111549006B (zh) * 2020-06-08 2023-03-28 杭州保安康生物技术有限公司 富含淀粉酶的复合酶的制备及其菌株和应用
CN111549020B (zh) * 2020-06-08 2023-03-28 杭州保安康生物技术有限公司 富含酸性果胶酶的复合酶的制备及其菌株和应用
CN112300945B (zh) * 2020-06-16 2022-09-27 青海民族大学 一种黑曲霉固态发酵产饲用复合酶制剂及其制备工艺
CN115747262A (zh) * 2021-09-03 2023-03-07 国投生物科技投资有限公司 利用小麦生产乙醇的方法
CN114081189B (zh) * 2021-10-09 2022-12-09 中国科学院微生物研究所 黑曲霉菌株酶制剂提高豌豆膳食纤维纯度的方法
CN114015580B (zh) * 2021-10-14 2023-08-08 吉林中粮生化有限公司 一种显著提高发酵饲料小肽含量的菌株及方法

Family Cites Families (4)

* Cited by examiner, † Cited by third party
Publication number Priority date Publication date Assignee Title
US3151983A (en) * 1961-05-09 1964-10-06 Nopco Chem Co Animal feed
SU489787A1 (ru) * 1974-01-02 1975-10-30 Внии Биосинтеза Белковых Веществ Главного Управления Микробиологической Промышленности Штамп плесневого гриба 22/12-продуцент карбогидразного комплекса ферментов
SU988867A1 (ru) * 1981-07-23 1983-01-15 Всесоюзный Научно-Исследовательский Институт Продуктов Брожения Способ приготовлени посевного материала дл культивировани плесневых грибов вида aSpeRGILLUS аUтамоRI,продуцирующих глюкоамилазу (его варианты)
FR2701487B1 (fr) 1993-02-10 1995-03-31 Agronomique Inst Nat Rech Réacteur pour conduire de façon stérile des procédés de fermentation d'un produit à l'état solide.

Also Published As

Publication number Publication date
DK1022329T3 (da) 2005-09-05
EP1022329B1 (fr) 2005-05-04
FR2788782A1 (fr) 2000-07-28
CA2361264A1 (en) 2000-07-27
ATE294852T1 (de) 2005-05-15
CN1337998A (zh) 2002-02-27
MA25276A1 (fr) 2001-10-01
AU775240B2 (en) 2004-07-22
ZA200107009B (en) 2002-07-31
AU3059800A (en) 2000-08-07
DE60019825T2 (de) 2006-04-27
TR200102156T2 (tr) 2002-05-21
PL350672A1 (en) 2003-01-27
US6667066B2 (en) 2003-12-23
JP2002534973A (ja) 2002-10-22
BRPI0007600B1 (pt) 2017-03-14
ES2244395T3 (es) 2005-12-16
HUP0104907A2 (hu) 2002-04-29
WO2000043496A1 (fr) 2000-07-27
CA2361264C (en) 2011-01-04
EP1022329A1 (fr) 2000-07-26
FR2788782B1 (fr) 2003-01-31
US20020037342A1 (en) 2002-03-28
PL201768B1 (pl) 2009-05-29
RU2246536C2 (ru) 2005-02-20
PT1022329E (pt) 2005-08-31
CN1203170C (zh) 2005-05-25
MXPA01007528A (es) 2003-03-27
DE60019825D1 (de) 2005-06-09
BR0007600A (pt) 2001-10-30

Similar Documents

Publication Publication Date Title
ID29922A (id) Produk multi-enzim dengan aktivitas glukoamilase, proteolitik dan ksilanase dan metode untuk memproduksinya dengan cara fermentasi kondisi padat dari kulit gandum dengan aspergillus niger
Gessesse et al. High-level xylanase production by an alkaliphilic Bacillus sp. by using solid-state fermentation
RU2001124277A (ru) Полиферментный продукт с глюкоамилазной, протеолитической и ксиланазной активностями, а также способ его получения путем твердофазного сбраживания пшеничных отрубей с помощью Aspergillus niger
ES511504A0 (es) "procedimiento para preparar un producto de enzima, para eliminacion de ramificaciones, capaz de hidrolizar enlaces alfa-1,6-glicosidicos".
Alazard et al. Comparative study of amylolytic enzymes production by Aspergillus niger in liquid and solid-state cultivation
MX173082B (es) Un procedimiento para macerar cereales con una nueva preparacion de enzima y producto de licor de maceracion
YU61799A (sh) Postupak za fermentacionu proizvodnju d-pantotenske kiseline pomoću pojačanja pan d-gena u mikroorganizmima
Gordon et al. Extracellular pectin lyase produced by Neocallimastix sp. LM1, a rumen anaerobic fungus
Han et al. Amylolysis of raw corn by Aspergillus niger for simultaneous ethanol fermentation
CN106635843B (zh) 发酵培养基、菌糠发酵菌剂及发酵方法
Conrad Enzymatic hydrolysis of xylans I. A high xylanase and β-xylosidase producing strain of aspergillus niger
Haska et al. Effect of cellulase addition on hydrolysis of sago starch granules by raw starch digesting amylase from Penicillium brunneum No. 24
Paszczyński et al. The influence of proteases on the activity of glucoamylase from Aspergillus niger C
Murty et al. Purification and properties of an extra cellular xylanase enzyme of Clostridium strain SAIV
Khare et al. Production of biodegradative enzymes by some thermophilic fungi of Agaricus bisporus compost
Mahmoud Different factors affecting growth and amylase production by fungi inhabiting poultry feeds
SU1252336A1 (ru) Штамм @ @ АТ-490 - продуцент целлюлаз
KOBAYASHI et al. Enzymatic Hydrolysis of" Polydextrose"
Rabinovitch et al. Thermostability of intracellular cellobiase of Thermomonospora sp.
RU96120611A (ru) Способ получения лимонной кислоты
JPH02255620A (ja) 抗白癬菌剤およびその製造方法、利用方法
Shaker et al. Optimization of the composition of the nutrient medium for cellulase and protein biosynthesis by thermophilic Aspergillus spp.
BG51306A1 (en) Method for preparation of enzyme preparation having leading xylanase activity
FR2353232A2 (fr) Production d'aliments a haute teneur en proteines par fermentation directe de produits broyes riches en amidon
D’Souza et al. Influence of pH on the formation and location of β-glucosidase in Aspergillus terreus grown on cellulose